I call this "adaptive dependency" (as among Buddhists' "dependent co-arising", or evolutionists/Darwinians' "co-evolution").

For 10+ years I have been learning about dependencies on one 100-acre area. Dependencies are far and beyond what I first thought when I started to live off/by (and for) the land (and what live on it). External climate is obvious. Micro-organisms are not. Migration (many birds, lizards, insects, and fungi are migratory) may bring in interesting (random?) changes.

I had since given up on "controlling"/"regulating" (my dream) dependencies (your "multi-symbiosis") and am now learning to watch to see what each day unfolds on the land.

Example: yesterday, a fresh water turtle came up from the dam, dug a little hole in the veggie patch about 10m from the water, laid 4 eggs (1/4 the size of chicken eggs, white soft shell, more elongated), she covered the hole, dragged herself back to the dam. I looked up the Net and read -- the eggs should hatch out in 1-3 months! (No one really knows. Chicken 20-22 days, duck 28-32 days, goose 40 days, depending on...?)

I think we know less about biology than we do know about physics ;-)